SGI Southern Appalachian coordinator Zach Irick leading vegetation monitoring in a bog in Marion County, Tennessee. The dominant plant around the boat is the rare Horsetail Spikerush (Eleocharis equisetoides), known to persist in Tennessee only at this site. Photo by Quentin Miller.

 SGI’s southern Appalachian coordinator Zach Irick monitoring a population of the federally listed endangered Green Pitcherplant (Sarracenia oreophila) at Dulaney Bog in North Carolina. Photo by Theo Witsell.

Specimen of the globally rare Small-headed Pipewort (Eriocaulon koernickianum) in a seepage wetland within a high-quality remnant tallgrass prairie in the Arkansas Valley of central Arkansas. This population represents a previously unknown site for this species and the third known from this rare habitat type. Photo by Theo Witsell.

 SGI's Southern Appalachian Grasslands Coordinator Zach Irick exploring a small bog at the base of a cliff deep in the backcountry of Cumberland Gap National Historical Park. Photo by Alaina Krakowiak.

Specimen of the globally rare Small-headed Pipewort (Eriocaulon koernickianum) from a seepage wetland within a high-quality remnant tallgrass prairie in the Arkansas Valley of central Arkansas. This population represents a previously unknown site for this species and the third known from this rare habitat type. Photo by Theo Witsell.

Snow-on-the-Prairie (Euphorbia bicolor) and Bottlebrush Blazing-star (Liatris punctata var. mucronata) in a high-quality remnant of a rare Limestone Prairie in the Red River Valley near Idabel, Oklahoma. Photo by Theo Witsell.

Grass-pink Orchid (Calopogon tuberosus) in a high-quality Longleaf Pine Savanna at the Greenwood Plantation in the Red Hill Region of southern Georgia. Photo by Theo Witsell.

 Rare remnant of an undescribed and globally rare Ozark Sand Grassland. This grassland type is known only from a small part of Izard County Arkansas in valleys of south-flowing tributaries of the White River. It supports many rare and uncommon plant species and the Ozark Pocket Gopher (Geomys bursarius ozarkensis), a globally rare subspecies known only from the southern third of Izard County and nowhere else on Earth. Photo by Theo Witsell.

A male Diana Fritillary (Speyeria diana), a butterfly of global conservation concern, in a restored and fire-managed shale barrens and woodland complex in the eastern Ouachita Mountains. Photo by Theo Witsell.

A specimen of the wonderfully named Bog Cheeto or Red-hot-poker (Senega lutea) in a high-quality Longleaf Pine Savanna at the Greenwood Plantation in the Red Hill Region of southern Georgia. Photo by Theo Witsell.

SGI's Central Appalachian Grasslands Coordinator Alaina Krakowiak coring an Eastern red cedar tree (Juniperus virginiana) that is nearly 200 years old. Photo by Laura Boggess.

SGI’s Eastern Region Biologist Jared Gorrell with a Prairie Kingsnake (Lampropeltis calligaster) in a fire-maintained woodland in the Ouachita Mountains. Photo by Theo Witsell.

A reference quality Ouachita Mountain Dry Sandstone Woodland at the Camp Robinson Special Use Area in central Arkansas. This site, with a 100-year prescribed fire history and highly diverse plant community, can be used to assess the quality and restoration progress of other sites across the region. Photo by Theo Witsell.

SGI's Central Appalachian Grasslands Coordinator Alaina Krakowiak documenting a population of Alabama lip fern (Myriopteris alabamensis) on a cliff face at Cumberland Gap National Historical Park. Photo by Laura Boggess.

A rare groundwater seep within a remnant Tallgrass Prairie. This rare combination of conditions supports several species of conservation concern including Small-headed Pipewort (Eriocaulon koernickianum).

SGI's Central Appalachian Grasslands Coordinator Alaina Krakowiak standing on a narrow ledge on cliff face at Cumberland Gap National Historical Park, where she surveyed for rare plant species by rappelling. Photo by Laura Boggess.

 Restoration technician Erin Gamst documenting prairie willow (Salix humilis) while sampling a vegetation plot in a degraded grassland remnant at Fredericksburg and Spotsylvania National Military Park. Photo by Alaina Krakowiak.

Herbarium specimen of Showy Beardtongue (Penstemon cobaea) from a high-quality Blackland Prairie remnant in the West Gulf Coastal Plain of Southwest Arkansas. Photo: ANHC.

 Herbarium specimen of Plantain-leaf Sunflower (Helianthus occidentalis subsp. plantagineus) from a rare hill prairie remnant in the Arkansas Valley Ecoregion. Photo: ANHC.

Herbarium specimen of Prairie Parsley (Polytaenia nuttallii) from a remnant shale barrens in the Ouachita Mountains. Photo: ANHC.

Herbarium specimen of Sticky Goldenrod (Solidago racemosa) from riverscour barrens in Big South Fork National River and Recreation Area on the Cumberland Plateau of Tennessee. Photo: APSC.

 Herbarium specimen of Virginia Bunchflower (Melanthium virginicum) from a rare seasonally-wet shale savanna in the Ouachita Mountains. Photo: ANHC.

 Herbarium specimen of Greethread (Thelesperma filifolium) from a remnant shale barrens in the Ouachita Mountains. Photo: ANHC.

 Herbarium specimen of Smoketree (Cotinus obovatus) from dolomite glades and barrens in the White River Hills Ecoregion of the Ozarks, collected near the Arkansas-Missouri border. Photo: ANHC.

 Herbarium specimen of Culver’s Root (Veronicastrum virginicum) from a pine-oak woodland in the Ozark Highlands. Photo: ANHC.

 Tennessee Natural Heritage Program Ecologist Adam Dattilo sampling a Cumberland Plateau Sandstone Glade and Barrens community in a Tennessee Valley Authority powerline right-of-way at Laurel Snow State Natural Area in Tennessee. This work was part of a collaborative study by SGI, the Tennessee Valley Authority, and the Electric Power Research Institute. Photo by Theo Witsell.

SGI Field Technician Brittney Georgic sampling remnant grassland vegetation in a Tennessee Valley Authority powerline right-of-way on the Cumberland Plateau of Tennessee. This work was part of a collaborative study by SGI, the Tennessee Valley Authority, and the Electric Power Research Institute. Photo by Theo Witsell.

 SGI Field Technician Brittney Georgic inspecting a population of Appalachian Milkwort (Senega curtissii) in a Tennessee Valley Authority powerline right-of-way on Bledsoe State Forest in Tennessee. This work was part of a collaborative study by SGI, the Tennessee Valley Authority, and the Electric Power Research Institute. Photo by Theo Witsell.

SGI Southern Appalachian Grasslands Coordinator Zach Irick in SGI’s Leatherflower (Clematis) Research Garden at Austin Peay State Univeristy. The garden allows ex situ cultivation of more than 75 different accessions of leatherflowers from across the Southeast. Ongoing research on the genus by SGI staff and partners has already resulted in the description of several new species from southeastern grasslands, with several more to come. Photo by Theo Witsell.

 High-quality grassland remnant in a powerline corridor at Cumberland Trail State Park in Cumberland County, TN. This grassland flora represents the natural ground cover of open savannas found historically in this region of the Cumberland Plateau. Today, these natural communities are rare due to impacts from past land use and encroachment by closed forest in the absence of fire and native herbivores. Photo by Theo Witsell.

 High-quality grassland remnant in a powerline corridor at Cumberland Trail State Park in Cumberland County, TN. This grassland flora represents the natural ground cover of open savannas found historically in this region of the Cumberland Plateau. Today, these natural communities are rare due to impacts from past land use and encroachment by closed forest in the absence of fire and native herbivores. Photo by Theo Witsell.

 High-quality grassland remnant in a powerline corridor at Cumberland Trail State Park in Cumberland County, TN. This grassland flora represents the natural ground cover of open savannas found historically in this region of the Cumberland Plateau. Today, these natural communities are rare due to impacts from past land use and encroachment by closed forest in the absence of fire and native herbivores. Photo by Theo Witsell.

 High-quality grassland remnant in a powerline corridor at Cumberland Trail State Park in Cumberland County, TN. This grassland flora represents the natural ground cover of open savannas found historically in this region of the Cumberland Plateau. Today, these natural communities are rare due to impacts from past land use and encroachment by closed forest in the absence of fire and native herbivores. Photo by Theo Witsell.

 SGI’s Western Region Biologist Allison Wilson sampling a high-quality riverscour barrens community on the Maumelle River in the eastern Ouachita Mountains of Arkansas. Photo by Virginia McDaniel.

 High-quality grassland remnant in a powerline corridor at Cumberland Trail State Park in Cumberland County, TN. This grassland flora represents the natural ground cover of open savannas found historically in this region of the Cumberland Plateau. Today, these natural communities are rare due to impacts from past land use and encroachment by closed forest in the absence of fire and native herbivores. Photo by Theo Witsell.

Cumberland Rosemary (Conradina verticillata) growing in high-quality riverscour barrens along the Clear Fork River within the Big South Fork National River and Recreation Area in Tennessee. This species is listed as Threatened under the federal Endangered Species Act and is found only in riverscour habitat on the Cumberland Plateau of Kentucky and Tennessee. Photo by Theo Witsell.

 High-quality grassland remnant in a powerline corridor at Cumberland Trail State Park in Cumberland County, TN. This grassland flora represents the natural ground cover of open savannas found historically in this region of the Cumberland Plateau. Today, these natural communities are rare due to impacts from past land use and encroachment by closed forest in the absence of fire and native herbivores. Photo by Theo Witsell.

High-quality riverscour barrens along the Clear Fork River within the Big South Fork National River and Recreation Area in Tennessee. Photo by Theo Witsell.

Yellow-fringed Orchid (Platanthera ciliaris) growing in a grassland remnant in a powerline corridor at Cumberland Trail State Park in Cumberland County, TN. Photo by Theo Witsell.

 Beautiful Barbara’s-buttons (Marshallia pulchra), a globally-rare plant listed as Endangered in both Kentucky and Tennessee, growing in high-quality riverscour barrens along the Clear Fork River within the Big South Fork National River and Recreation Area in Tennessee. Photo by Theo Witsell.

 Yellow Nodding Ladies’-tresses (Spiranthes ochroleuca) in a powerline right-of-way at Big South Fork National River and Recreation Area in Tennessee. This is one of many species of sun-loving savanna and open woodland plants hanging on in open rights-of-way after surrounding woodlands have become too dense to support them. Photo by Theo Witsell.

 SGI Chief Conservation Officer Theo Witsell and Executive Director Dwayne Estes with a large haul of plant specimens from riverscour habitats on the South Fourche LaFave River in the Ouachita Mountains of Arkansas. Photo by Todd Crabtree.

Zigzag Bladderwort (Utricularia subulata) in a seep in a powerline right-of-way at North Chickamauga Creek Wildlife Management Area in Tennessee. This carnivorous plant is one of many species of sun-loving savanna and open woodland plants hanging on in open rights-of-way after surrounding woodlands have become too dense to support them. Photo by Theo Witsell.

SGI field staff in a plant identification training at the Camp Robinson Special Use Area in Central Arkansas. Photo by Theo Witsell.

SGI’s Western Region Biologist Allison Wilson (right) and volunteer Virginia McDaniel (left) sampling an excellent-quality riverscour barrens on the Maumelle River in the eastern Ouachita Mountains. Photo by Ben Thesing.

Tennessee Natural Heritage Program Ecologist Adam Dattilo sampling grassland flora in a Tennessee Valley Authority powerline right-of-way. This work was part of a collaborative study by SGI, the Tennessee Valley Authority, and the Electric Power Research Institute. Photo by Theo Witsell.

 SGI’s western field team sampling the plant community in a high-quality saline barrens and savanna complex at Kingsland Prairie Natural Area in the West Gulf Coastal Plain of Arkansas. Photo by Theo Witsell.

 SGI’s Southern Appalachian Grassland Coordinator Zach Irick (center) and volunteer Ben Benton (foreground) conducting surveys of riverscour habitats along the Clear Fork River within the Big South Fork National River and Recreation Area in Tennessee. Photo by Theo Witsell.

SGI’s western field team using iNaturalist to document the occurrence of Oneflower Hawthorn (Crataegus uniflora) in a rare sand grassland in the Ozark Highlands. Photo by Theo Witsell.

SGI’s Southern Appalachian Grassland Coordinator Zach Irick cooling off while traversing the narrow gorge of the Clear Fork River during riverscour surveys of the Big South Fork National River and Recreation Area in Tennessee. Portions of the gorge are lined with cliffs or house-sized boulders making the river channel the safest path. Photo by Theo Witsell.

 SGI’s Eastern Region Ecologist Vero Tessier documenting an occurrence of Buffalo Clover (Trifolium reflexum) in a restored pine-oak woodland in the eastern Ouachita Mountains. Photo by Theo Witsell.

SGI’s Southern Appalachian Grassland Coordinator Zach Irick with a large population of Virginia Sweetspire (Spiraea virginiana), growing in high-quality riverscour barrens along the Clear Fork River within the Big South Fork National River and Recreation Area in Tennessee. This species is listed as Threatened under the federal Endangered Species Act and is found only in riverscour habitat. Photo by Theo Witsell.

 SGI’s Eastern Region Ecologist Vero Tessier documenting an occurrence of Buffalo Clover (Trifolium reflexum) in a restored pine-oak woodland in the eastern Ouachita Mountains. Photo by Theo Witsell.

SGI’s Western Region Biologist Allison Wilson sampling a high-quality riverscour barrens community on the Maumelle River in the eastern Ouachita Mountains of Arkansas. Photo by Virginia McDaniel.

SGI field staff training in vegetation plot sampling in an old-growth post oak savanna/woodland complex at the Camp Robinson Special Use Area in Central Arkansas. Photo by Theo Witsell.

SGI field staff training in vegetation plot sampling in an old-growth post oak savanna/woodland complex at the Camp Robinson Special Use Area in Central Arkansas. Photo by Theo Witsell.

SGI field staff training in vegetation plot sampling in a high-quality seasonally wet prairie remnant at the Camp Robinson Special Use Area in Central Arkansas. Photo by Theo Witsell.